There are two life history landmarks that can be used to define the terminal period in individual Drosophila melanogaster females: the cessation of daily oviposition, which defines the start of the retired stage, and final oviposition, which defines the start of post-ovipository survival. The terminal period is a substantial component of D. melanogaster life history. Analysis of published data on the daily fecundity and survival of 3971 individually maintained, mated female flies reveals that the terminal period is far more variable within populations than other life history components, including total adult life span. It has been reported that there is a negative correlation between fecundity and duration of the terminal state in recently collected wild stocks. Here I show that the negative correlation occurs in multiple inbred and outbred lab-adapted populations as well. In terms of proportion of adult life, lower fecundity flies spend on average twice as much time in the terminal stage as higher fecundity flies from the same population. Both high and low fecundity flies experience end-of-life plateaus in mortality, with the former exhibiting higher plateau levels. The negative correlation between fecundity and terminal survival is of sufficient magnitude to create heterogeneity among the oldest old: the final 10% of survivors are predominately flies with a history of high fecundity, but about one in five is a low fecundity fly with long terminal stage.

译文

:有两个生活史上的里程碑可用来定义果蝇雌性个体的终末期:停止每日排卵(这决定了退休阶段的开始)和最终排卵(这决定了排卵后生存的开始) 。末期是黑腹果蝇生活史的重要组成部分。对已发表的3971只个体交配的雌性苍蝇的每日繁殖力和存活率的公开数据进行的分析表明,与其他生命史组成部分(包括总成年寿命)相比,种群内的终末期变化更大。据报道,在最近采集的野生种群中,繁殖力与绝热状态持续时间之间呈负相关。在这里,我表明负相关也发生在多个近交和近交实验室适应人群中。就成年生活的比例而言,繁殖力低下的苍蝇在末期平均花费的时间是同一种群的繁殖力高的苍蝇的两倍。高繁殖力和低繁殖力的苍蝇都经历了生命的停滞高原,而前者表现出较高的高原水平。繁殖力与最终生存之间的负相关程度足以在最老的老年人中造成异质性:最后10%的幸存者主要是具有高繁殖力历史的果蝇,但大约五分之一是终末期长的低繁殖力果蝇。

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