Viruses cause significant mortality of marine microorganisms; however, their role in shaping the composition of microbial assemblages has not been fully elucidated. Because viruses may form lysogenic relationships with their hosts, temperate viruses may influence bacterial assemblage structures through direct lysis of hosts when induced by environmental stimuli or by homoimmunity (i.e., immunity to closely related viruses). We investigated the components of bacterioplankton assemblages that bore prophage using the lysogenic induction agent mitomycin C. Seawater was collected at two locations (the San Pedro Ocean Time Series Station and in the Santa Barbara Channel) in the Southern California Borderland and amended with mitomycin C. After 24-h incubation, the community structure of bacterioplankton was compared with unamended controls using automated rRNA intergenic spacer analysis. The addition of mitomycin C to seawater had effects on the community structure of bacterioplankton, stimulating detectable overall diversity and richness of fingerprints and causing the assemblages within incubations to become different to control assemblages. Most negatively impacted operational taxonomic units (OTU) in mitomycin C-amended incubations individually comprised a large fraction of total amplified DNA in initial seawater (5.3-23.3% of amplified DNA fluorescence) fingerprints, and data suggest that these include organisms putatively classified as members of the gamma-Proteobacteria, SAR11 cluster, and Synechococcus groups. The stimulation of assemblage richness by induction of lysogens, and the reduction in the contribution to total DNA of common OTU (and concomitant increase in rare OTU), suggests that temperate phage have the potential to strongly influence the diversity of bacterioplankton assemblages. Because lysogenic OTU may also be resistant to closely related lytic (i.e., free-living) viruses, the impact of lytic virioplankton on assemblages may only be pronounced transiently or when conditions causing lysogenic induction arise.

译文

:病毒导致海洋微生物大量死亡;然而,它们在塑造微生物组合物的组成中的作用尚未完全阐明。因为病毒可能与其宿主形成溶源性关系,所以当受到环境刺激或同种免疫力(即对紧密相关的病毒的免疫力)诱导时,温带病毒可能通过直接裂解宿主来影响细菌的组装结构。我们调查了使用溶原性诱导剂丝裂霉素C进行噬菌体繁殖的浮游细菌集合体的组成部分。在南加利福尼亚边境地区的两个地点(圣佩德罗海洋时间序列站和圣塔芭芭拉海峡)收集了海水,并用丝裂霉素C进行了修正。温育24小时后,使用自动rRNA基因间间隔子分析将浮游细菌的群落结构与未经修饰的对照进行比较。向海水中添加丝裂霉素C对浮游细菌的群落结构有影响,刺激了可检测的总体多样性和指纹的丰富性,并导致孵化过程中的组合变得与对照组合不同。在丝裂霉素C修饰的孵育中,最受负面影响的操作生物分类单位(OTU)分别包含初始海水指纹中总扩增DNA的很大一部分(占扩增DNA荧光的5.3%至23.3%),数据表明,这些微生物包括假定分类为成员的生物γ-变形杆菌,SAR11簇和Synechococcus组。通过溶菌原的刺激刺激丰富的集合体,以及减少普通OTU对总DNA的贡献(并伴随稀有OTU的增加),表明温和的噬菌体有可能强烈影响浮游细菌集合体的多样性。由于溶源性OTU也可能对紧密相关的溶菌(即自由生活)病毒具有抗性,因此溶溶性浮游生物对集合体的影响只能短暂地或在引起溶源性诱导的条件出现时才显露出来。

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