Sulphur deficiency in crops became an agricultural concern several decades ago, due to the decrease of S deposition and the atmospheric sulphur dioxide emissions released by industrial plants. Autophagy, which is a conserved mechanism for nutrient recycling in eukaryotes, is involved in nitrogen, iron, zinc and manganese remobilizations from the rosette to the seeds in Arabidopsis thaliana. Here, we have compared the role of autophagy in sulphur and nitrogen management at the whole plant level, performing concurrent labelling with 34S and 15N isotopes on atg5 mutants and control lines. We show that both 34S and 15N remobilizations from the rosette to the seeds are impaired in the atg5 mutants irrespective of salicylic acid accumulation and of sulphur nutrition. The comparison in each genotype of the partitions of 15N and 34S in the seeds (as % of the whole plant) indicates that the remobilization of 34S to the seeds was twice more efficient than that of 15N in both autophagy mutants and control lines under high S conditions, and also in control lines under low S conditions. This was different in the autophagy mutants grown under low S conditions. Under low S, the partition of 34S to their seeds was indeed not twice as high but similar to that of 15N. Such discrepancy shows that when sulphate availability is scarce, autophagy mutants display stronger defects for 34S remobilization relative to 15N remobilization than under high S conditions. It suggests, moreover, that autophagy mainly affects the transport of N-poor S-containing molecules and possibly sulphate.

译文

:几十年前,由于S沉积物的减少和工业工厂释放的大气中的二氧化硫排放,农作物中的硫缺乏成为农业关注的问题。自噬是真核生物中养分循环的一种保守机制,它参与了拟南芥中从花环到种子的氮,铁,锌和锰的固定。在这里,我们比较了自噬在整个植物水平上硫和氮管理中的作用,并在atg5突变体和对照品系上同时进行了34S和15N同位素标记。我们显示,无论是水杨酸积累还是硫营养,atg5突变体均会损害从莲座丛到种子的34S和15N迁移。种子中15N和34S分区的每种基因型的比较(占整个植物的百分比)表明,在高S下自噬突变体和对照品系中,将34S固定到种子上的效率是15N的两倍。以及低S条件下的控制线路中。这在低S条件下生长的自噬突变体中是不同的。在低S下,其种子中34S的分配确实不高,但与15N相似。这种差异表明,当缺乏硫酸盐时,相对于15N迁移,自噬突变体在34S迁移方面表现出比在高S条件下更强的缺陷。此外,这表明自噬主要影响含N贫S的分子和硫酸盐的转运。

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