Pseudomonas aeruginosa is a gram-negative opportunistic pathogen that is cytotoxic towards a variety of eukaryotic cells. To investigate the effect of this bacterium on monocyte, we infected human U937 cells with the P. aeruginosa strain in vitro. To explore the expression of Bcl-2 and Bax as well as caspase-3/9 activation in the apoptosis of human U937 cells induced by P. aeruginosa, Hoechst 33258 staining and Giemsa staining as well as Flow cytometry analysis were used to determine the rate of apoptosis, and the expressions of Bcl-2 and Bax were assayed by RT-PCR and Western blotting respectively. Bax protein conformation change was assayed by immunoprecipitation. Cytochrome c release was measured by Western blotting. Moreover, exposure of U937 cells to P. aeruginosa measured caspase-3/9 activity. It was found that the apoptosis of human U937 cells could be induced by Pseudomonas aeruginosa in a dose- and time-dependent manner. Also, there were a tendency of alterations with an increased expression level of Bax and a reduced expression level of Bcl-2, increased levels of cytochrome c release, and also with an increased activation of caspase-3/9 and Bax protein conformation change. For the evaluation of the role of caspases, caspase-3/9 inhibitors Z-DEVD-FMK and Z-LEHD-FMK respectively were used. The results were further confirmed by the observation that the caspase inhibitors Z-DEVD-FMK and Z-LEHD-FMK blocked P. aeruginosa-induced U937 apoptosis. It is concluded that P. aeruginosa can induce apoptosis with an up-regulated expression of Bax and a down-regulated expression of Bcl-2, which resulted in increased levels of cytochrome c release and increased caspase-3 and -9 in human U937 cells.

译文

铜绿假单胞菌是一种革兰氏阴性机会性病原体,对多种真核细胞具有细胞毒性。为了研究这种细菌对单核细胞的影响,我们在体外用铜绿假单胞菌菌株感染了人U937细胞。为探讨铜绿假单胞菌诱导人U937细胞凋亡中Bcl-2和Bax的表达以及caspase-3/9的激活,采用Hoechst 33258染色和Giemsa染色以及流式细胞仪分析测定凋亡率。用rt-pcr和Western blotting分别检测Bcl-2和Bax的表达。通过免疫沉淀测定Bax蛋白构象变化。通过蛋白质印迹法测量细胞色素c的释放。此外,暴露于铜绿假单胞菌的U937细胞可测量caspase-3/9活性。发现铜绿假单胞菌可以剂量和时间依赖性地诱导人U937细胞的凋亡。此外,随着Bax表达水平的增加和Bcl-2表达水平的降低,细胞色素c释放水平的增加以及caspase-3/9和Bax蛋白构象变化的增加,存在改变的趋势。为了评估半胱天冬酶的作用,分别使用caspase-3/9抑制剂Z-DEVD-FMK和Z-LEHD-FMK。观察结果进一步证实,caspase抑制剂Z-DEVD-FMK和Z-LEHD-FMK阻断了铜绿假单胞菌诱导的U937凋亡。结论铜绿假单胞菌可诱导凋亡,Bax表达上调,Bcl-2表达下调,导致人U937细胞细胞色素c释放水平升高,caspase-3和-9增加。

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