The majority of peripheral serotonin is stored in platelets, which secrete it on activation. Serotonin releases Weibel-Palade bodies (WPBs) and we asked whether absence of platelet serotonin affects neutrophil recruitment in inflammatory responses. Tryptophan hydroxylase (Tph)1–deficient mice, lacking non-neuronal serotonin, showed mild leukocytosis compared with wild-type (WT), primarily driven by an elevated neutrophil count. Despite this, 50% fewer leukocytes rolled on unstimulated mesenteric venous endothelium of Tph1(-/-) mice. The velocity of rolling leukocytes was higher in Tph1(-/-) mice, indicating fewer selectin-mediated interactions with endothelium. Stimulation of endothelium with histamine, a secretagogue of WPBs, or injection of serotonin normalized the rolling in Tph1(-/-) mice. Diminished rolling in Tph1(-/-) mice resulted in reduced firm adhesion of leukocytes after lipopolysaccharide treatment. Blocking platelet serotonin uptake with fluoxetine in WT mice reduced serum serotonin by > 80% and similarly reduced leukocyte rolling and adhesion. Four hours after inflammatory stimulation, neutrophil extravasation into lung, peritoneum, and skin wounds was reduced in Tph1(-/-) mice, whereas in vitro neutrophil chemotaxis was independent of serotonin. Survival of lipopolysaccharide-induced endotoxic shock was improved in Tph1(-/-) mice. In conclusion, platelet serotonin promotes the recruitment of neutrophils in acute inflammation, supporting an important role for platelet serotonin in innate immunity.

译文

:大多数外周血清素都储存在血小板中,血小板在激活时会分泌出来。血清素释放Weibel-Palade体(WPB),我们询问是否缺乏血小板血清素会影响中性粒细胞在炎症反应中的募集。缺乏非神经元5-羟色胺的色氨酸羟化酶(Tph)1缺陷小鼠与野生型(WT)相比,表现出轻度白细胞增多,主要是由于嗜中性白细胞计数升高所致。尽管如此,Tph1(-/-)小鼠未经刺激的肠系膜静脉内皮上滚动的白细胞减少了50%。 Tph1(-/-)小鼠中滚动白细胞的速度较高,表明较少的选择素介导的与内皮的相互作用。组胺刺激,外周血白蛋白的促分泌素或血清素注射刺激正常化Tph1(-/-)小鼠的滚动。 Tph1(-/-)小鼠的滚动减少导致脂多糖治疗后白细胞牢固粘附减少。氟西汀在野生型小鼠中阻断血小板对血清素的摄取,使血清素减少了80%以上,同样降低了白细胞的滚动和粘连。炎症刺激后四个小时,Tph1(-/-)小鼠中性粒细胞向肺,腹膜和皮肤伤口的渗出减少,而体外中性粒细胞趋化性独立于5-羟色胺。 Tph1(-/-)小鼠中脂多糖诱导的内毒素休克的存活率得到改善。总之,血小板5-羟色胺可促进中性粒细胞在急性炎症中的募集,支持血小板5-羟色胺在先天免疫中的重要作用。

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